Agroecosystem is a concurrence of highly complicated interaction such as crop and pest. An agroecosystem can be viewed as a subset of a conventional ecosystem, for example, to identify the pest distribution or seasonal population fluctuations etc. These are really important investigation to set the IPM strategy. With these study, now we focused on sophisticated interaction between pest and their parasite in genome and transcriptome level. We studied these interesting interaction between diamondback moth Plutella xylostella which major pests in cabbage and Diadegma fenestrale as a model endoparasitic wasp. Actually, D. fenestrale was NOT alone because it has symbiotic virus, D. fenestrale Ichnovirus (DfIV). We will deeply discussed these interaction between P. xylostella and D. fenestrale with DfIV.