Green mold disease caused by pathogenic Trichoderma spp. (such as T. aggressivum, T. harzianum, and T. pleurotum) is a major threat to the production of commercial mushrooms, including Agaricus bisporus, Pleurotus ostreatus, and Lentinula edodes. Trichoderma spp. act as aggressive competitors and mycoparasites that rapidly colonize substrates, secrete cell walldegrading enzymes, and release toxic metabolites, severely inhibiting host mycelial growth and yield. This review summarizes current knowledge on the diversity, epidemiology, and cellular interaction mechanisms of pathogenic Trichoderma spp. in cultivated mushrooms. We also evaluate the existing control strategies, including substrate pasteurization, chemical fungicides, biocontrol agents, and resistance breeding. Future directions for advancing eco-friendly integrated pest management (IPM) frameworks for sustainable mushroom cultivation are discussed.

ABSTRACT
INTRODUCTION
Taxonomy and Causal Agents
Genetic Diversity
Pathogenic Mechanisms and Mycoparasitism
    1. Enzymatic Degradation Trichoderma secretes a complex suite of Cell Wall Degrading Enzymes (CWDEs) (Kredics et al., 2021):
    2. Antibiosis and Mycotoxins
Symptoms and Economic Impact
    1. Agaricus bisporus (Button Mushroom)
    2. Pleurotus spp. (Oyster Mushroom)
    3. Lentinula edodes (Shiitake)
Morphological Characteristics
Diagnostic Methodologies
Integrated Disease Management (IDM) Strategies
    1. Hygiene and Environmental Sanitation
    2. Biological Control
    3. Chemical Control and Resistance Challenges
Conclusions and Future Research Directions
ACKNOWLEDGEMENTS
REFERENCES

• Woo-Sik Jo(Gyeongbuk Province Agricultural Technology Administration, Division of Agricultural Food and Environmental Research Daegu, 41404, Korea) Corresponding author