Insect pests on cruciferous crops and their natural enemies were surveyed during 2009~2011 in Daegwallyeong highland region which has been famous for summer Kimchi cabbage production in Korea since 1970s. Totally 15 insect pests have been reported in cabbage field in Daegwallyeong. Diamondback moth (DBM, Plutella xylostella) imported cabbage worm (Artogeia rapae L.), cabbage armyworm (CAW, Mamestra brassicae L.), cabbage looper (CL, Trichoplusia ni), cabbage sawfly (Athalia rosae ruficornis Jakovlev), and turnip aphid (Liphapis pseudobrassicae (Davis)) were the dominant pest species among the 15 species. For monitoring as well as controlling insects with weak flight, yellow sticky cards (10×15㎝) were used in cabbage fields. The sticky cards were hanged on plastic stick and placed at a 2-5 m distance within cabbage field (1,000㎡). Sex pheromone traps were employed for monitoring and controlling insects with strong flight. The survey result showed that there was significantly reduced pest occurrences in treated, compared to untreated condition. Similarly, The results from the sex pheromone traps were coincident with those of sticky trap. DBM, CAW and CL were found less in the treated than untreated fields; by 67.5%, 70.6% and 44.0%, respectively. Although the control efficacy of such traps was less than that of chemical spray, the use of sticky card trap combined with sex pheromone trap could be useful management tools for controlling insect pests in cabbage fields.

같은 속의 복숭아순나방(Grapholita molesta)과 복숭아순나방붙이(G. dimorpha)는 기주범위와 야외에서의 성충 발생 양상이 유사하다. 또한 두 종은 성페로몬 성분으로 cis-8-dodecenyl acetate (Z8-12:Ac)와 trans-8-dodenceyl acetate (E8-12:Ac)을 공통으로 사용하고 있는데, 이 두 성분으로 구성된 상용 성페로몬 유인제에 두 종이 혼합되어 유인된다. 본 연구는 복숭아순나방붙이 트랩에 복숭아순나방이 혼합유인되는 현상을 줄이기 위해 성페로몬 미량성분의 효과를 분석했다. 복숭아순나방붙이 성페로몬인 Z8-12:Ac/E8-12:Ac = 85/15에 cis-8-tetradecenyl acetate(Z8-14:Ac)를 첨가했을 때, 복숭아순나방붙이의 유인은 유지되면서 복숭아순나방의 유인은 유의하게 억제되었다. Dodecyl acetate (12:Ac)와 tetradecyl acetate (14:Ac)의 첨가는 두 종의 유인력에 영향을 주지 않았다. 한편 Z8-12:Ac/E8-12:Ac/Z8-14:Ac = 85/15/10의 세 성분 성페로몬에 trans-8-tetradecenyl acetate (E8-14:Ac)를 첨가했을 때, 복숭아순나방붙이의 유인이 오히려 유의하게 억제되었다. 이상의 결과로부터 Z8-12:Ac/E8-12:Ac/Z8-14:Ac = 85/15/10의 조성이 개선된 복숭아순나방붙이의 성페로몬으로 제시되었다.

배나무에서 가루깍지벌레의 합성 성페로몬을 이용하여 수컷 성충의 발생 양상과 교미교란 효과 검토를 통해 효과적인 방제 정보를 제공하고자 본 시험을 수행하였다. 가루깍지벌레 성페로몬(2,6-dimethyl-1,5-heptadien-3-yl acetate)이 3 mg과 1.5 mg씩 침적된 고무 캡으로 수컷 유인수를 조사하였는데, 총포획수는 1.5 mg을 침적시킨 트랩에 1,021마리, 3 mg을 침적시킨 미끼에는 1,431마리의 수컷 성충이 포획되어 3 mg을 침적시킨 미끼에 더 많이 유인되었다. 황색과 백색 트랩에 의한 포획량의 차이는 황색트랩에 평균 포획수가 약간 더 많았지만 유의성은 없었다. 가루깍지벌레의 수컷 성충은 2012년에 6월부터 10월까지 포획되었는데, 6월 중순에 가장 많은 수가 포획되었고, 8월 상순, 9월 하순에도 발생 peak를 보였다. 2013년에도 6월부터 10월까지 포획되었는데, 6월 중순에 가장 높은 발생 최성기를 보였고, 7월 하순, 9월 하순에도 발생 최성기를 보였다. 가루깍지벌레 성페로몬을 이용한 교미교란제를 10 a당 450, 900, 및 1,350 mg의 농도로 과수원에 설치한 경우, 수컷 성충의 트랩 유인저해율은 각각 17.7, 65.3, 62.9%였다. 한편, 10a당 450, 900, 및 1,350mg 처리한 시험구에서 수확기에 피해과율이 각각 3.4, 2.9, 4.8%를 보인 반면, 교미교란제를 처리하지 않은 시험구에서는 9.5%였다.

A male specimen of Lymantria albescens (called as Okinawa gypsy moth) was captured in Busan, by sex-pheromone trap for Asian Gypsy Moth (AGM) (7R,8S)-cis-7,8-epoxy-2- methyloctadecane [(+)-disparlure]. Up to now, this species is distributed only in Ryukyu Islands of Japan including Ishigaki and Okinawa. The male of Okinawa gypsy moth might be attracted to AGM pheromone trap. If L. albescens is occurred in Korea, more many male individuals must be captured in pheromone trap. Therefore, we considered that the individual might be imported from Japan by inanimate pathway. Although it is high probability that L. albescens might be imported from Okinawa, it is important to a survey on an invasive pathway of the species in a view point of quarantine inspection. Through this presentation, we provided a detection method on Lymantria species using DNA barcoding. On the basis of this study, we will conduct on an invasive pathway and inhabitation possibility.

The seasonal occurrence of two Tortricinae species, Adoxophyes honmai (Yasuda) and Homona magnanima Diakonoff, was monitored at persimmon orchard in Changwon city, Gyeongnam, using sex attractant traps in 2012 and 2013. During the monitoring, a number of Neocalyptis angustilineata (Walsingham) was attracted to sex pheromone traps of H. magnanima. For A. honmai, a blend of Z9-14Ac, Z11-14Ac, E11-14Ac, 10me-12Ac with the ration of 314, 623, 55, and 9 ㎍ respectively, was used. A blend of Z11-14Ac and Z9-12Ac in the ratio of 900+100 ㎍ was used for H. magnanima. A 1:1 blend of Z11-14Ac and Z9-12Ac (500+500 ㎍) was also used for N. angustilineata. The two-years’ monitoring revealed that A. honmai and H. magnanima occurred four times a year, and N. angustilineata did three times. The peak occurrence of each generation for A. honmai was the 2nd or 3rd week of May, the 1st to 2nd week of July, and 5th week of July to 1st week of August, and 2nd to 4th week of September for the 1st, 2nd, 3rd, and 4th generation, respectively. The peak occurrence of each generation for H. magnanima was the 3rd to 4th week of May, the 2nd to 3rd week of July, and 4th week of August to 1st week of September, and 4th week of September to 3rd week of October for the 1st, 2nd, 3rd, and 4th generation, respectively. N. angustilineata showed clear three peaks of the 5thweek of May to 1st week of June, from 4th to 5th week of July, and 1st to 3rd week of September for the 1st, 2nd, and 3rd generation, respectively. A significantly higher number of N. angustilineata was attracted to sex pheromone traps of H. magnanima for the two years than H. magnanima was. H. magnanima was attracted only to the 9:1 blending ratio. However, N. angustilineata was attracted equally to both of the blending ratios.

Apolygus spinolae (Meyer-Dür) (Heteroptera: Miridae) is an important pest of fruit and tea trees in Korea and Japan. Analyses of extracts of metathoracic scent glands revealed that those of female bugs contained hexyl butyrate, (E)-2-hexenyl butyrate, and (E)-4-oxo-2-hexenal in a ratio of 20:100:7. The glands of males contained the same three compounds, but the ratio of the components was quite different, with hexyl butyrate being the most abundant. Field trapping tests with various blends of the synthetic compounds dispensed from high-density polyethylene tubes showed that (E)-2-hexenyl butyrate and (E)-4-oxo-2-hexenal were essential for attraction of male A. spinolae, and catches with a wide range of ratios of these two compounds did not differ significantly. However, adding hexyl butyrate at 50% or more of the (E)-2-hexenyl butyrate to the binary blend strongly inhibited attraction of males. Trap catches increased with increasing amounts of a 10:1 blend of (E)-2-hexenyl butyrate and (E)-4-oxo-2-hexenal from 0.011 to 11 mg loaded into the tube. Catches of males in traps baited with lures containing 1.1 mg of the binary blend were not significantly different from catches in traps baited with live virgin females.

The effect of increased carbon dioxide concentration in atmosphere was examined on the pheromone system of Helicoverpa armigera reared from egg stage to adult in three room. Two of three room (2×2×2 m) were treated with carbon dioxide gas as 600 ppm and 1,000 ppm, respectively. Mean of carbon dioxide concentration was 429.1 ppm in the control, 603.3 ppm for 600 ppm, and 1011.5 ppm for 1,000 ppm during experiment. Electroantenograph (EAG) test was conducted on 3-d-old male adults with air, hexane, and a series of their sex pheromone component, Z11-16Al, from 0.01 to 100 ng. The result was that male EAG responses of 600 and 1,000 ppm were 30.3% lower than that of control room. Production of Z11-16:Al was examined on about twenty 2-d-old virgin females. Carbon dioxide increases did not show a statistically significant difference. However, higher amount of sex pheromone was produced in females of 600 and 1,000 ppm. So, This experiment was replicated with different population reared again. The amount of the sex pheromone per female was 108.9 and 118.1 ng in control room, 139.8 and 141.8 ng in 600 ppm room, and 124.6 and 125.8 ng in 1,000 ppm room.

Prior to 1990s, most farmers customarily sprayed the agrochemicals that only just recommended by association or pesticide dealers, regardless of the monitoring for the disease incidences and pest occurrences. In 1993, agrochemicals sprayed in apple orchards a year had been done about 15 ~16 times. As founded the Apple Research Station (ARS) in 1991, a study for the construction of Integrated Pest Managements (IPMs) were actively done during 1990s and the IPM was paved the way to control on apple diseases and pests during the 2000s. The co-work between ARS and Seoul National University during 1996 to 2001 for the monitoring and development of IPM strategies using sex pheromone made easy understanding for IPM and occurrence monitoring results by showing the pests to apple farmers. Through the 2000s, IPM pilot project for the distribution of IPM technique utilizing sex pheromone trap and mating disrupter to the field has endeavored. Currently, sex pheromone trap plays an important role for monitoring of the major apple pests and the quarantine pest, C. sasakii on apple. Moreover, sex pheromonebased mating disruptor has been used almost all organic apple orchards for the control of one of major fruit-infesting pest, G. molesta and C. sasakii. Henceforth, the application of sex pheromone trap for monitoring as well as the active expansion and provision of mating disruptor for the control is needed. To this, first of all, it is very important to reduce prices by developing the techniques than other things. Second, reinforced education for monitoring staffs must be done continuously to instruct and to consult the farmers with the monitoring results to give guidance. Third, companies competing in good faith will improve and maintain their product quality.

비래해충인 멸강나방 성충의 발생을 실시간으로 예찰하기 위하여, 성페로몬 트랩에 원격감지장치를 설치한 트랩의 운용 효율성을 검정하였다. 원격감지트랩은 콘트랩 기반에 곤충의 유입을 광차단으로 감지하는 센서부, 신호전송부, 태양열기반의 전력공급부, 트랩 지지대, 웹페이지 기반의 자료수집부로 구성되었다. 연구과정에서 멸강나방 수컷 성충의 밤 시간대 포획의 일일주기성을 이용한 신호감지 시간대 축소, 신호발생 프로그램 조절, 1시간 간격의 전송간격, 신호전송 프로그램 변형 등을 통해 포획 나방수와 감지신호 발생 수의 격차를 줄여 최종적으로 신호 정확도를 92% 이상으로 개선하였다. 또한 실제 멸강나방 성충의 발생양상과 원격감지트랩에서의 신호 발생양상의 상관계수가 0.98이상으로 원격감지트랩에서의 신호 발생이 멸강나방 발생 양상을 정확하게 반영하는 것으로 나타났다. 2011~2012년 멸강나방은 연중 여러 번 발생되는 것이 관찰되었는데, 모두 짧은 기간 동안 발생되어 가파른 피크가 특징이었다.

The oriental fruit moth (Grapholita molesta) and the plum fruit moth (G. dimorpha) share the same major sex pheromone components, Z8-dodecenyl acetate (Z8-12Ac) and E8-dodecenyl acetate (E8-12Ac) with different ratio. However, these two congener male species were cross-attracted to the counter sex pheromone traps. For development of the specific monitoring lures, the minor sex pheromone components were added to the major components. G. molesta females emit two minor components of Z8-12OH and 12OH and G. dimorpha females emit four minor of 12Ac, 14Ac, Z8-14Ac, and E8-14Ac. For a specific monitoring lure of G. molesta, only Z8-12Ac major component attracted only G. molesta males, but did not any G. dimorpha. For a specific monitoring lure of G. dimorpha, the addition of Z8-14Ac to the major component (Z8-12Ac:E8-12Ac = 85:15) attracted G. dimorpha males with less than 5% G. molesta males. Other with components (12Ac, 14Ac, and E8-14Ac) was not effective in both trapping efficiency and selectivity.

The sex pheromone of Stathmopoda masinissa (Lepidoptera: Stathmopodidae) [감꼭지나방], an important pest of persimmon in Korea, was studied. Two pheromone components which known from Japanese population were detected in the gland extracts of females by mass spectral analysis of the gland extracts and comparisons of retention time with authentic compounds. Those were (E4,Z6)- 4,6-hexadecadienyl acetate (E4,Z6-16Ac) and (E4,Z6)-4,6-hexadecadien-1-ol (E4,Z6-16OH) in the ratio of 10:90. In a persimmon orchard, a single compound of E4,Z6-16Ac and E4,Z6-16OH was attractive to S. masinissa. However, the mixture of E4,Z6-16Ac and E4,Z6-16OH (1:1) caught significantly more males than each single component. In addition, the traps baited with E4,Z6-16Ac attracted significantly more Oedematopoda ignipicta (Lepidoptera: Stathmopodidae) [붉은꼬마꼭지나방] than control. These results suggest that both of the E4, Z6-16Ac and E4,Z6-16OH are needed to attract S. masinissa

The sex pheromone of Stathmopoda auriferella (Walker) (Lepidoptera: Stathmopodidae), an important pest of kiwifruit in Korea, was studied. Two candidate pheromone components detected in the gland extracts of females were identified as (E)-5-hexadecenyl acetate (E5-16:OAc) and (E)-5-hexadecenol (E5-16:OH) in a ratio of 75:25 by mass spectral analysis of natural pheromone components and dimethyldisulfide adducts, and retention index comparisons with synthetic standards. In the kiwifruit orchards, E5-16:OAc alone was attractive to S. auriferella males and caught significantly more males than live virgin females. However, addition of E5-16:OH strongly inhibited attraction to E5-16:OAc. These results suggest that the major component of the female-produced sex pheromone of S. auriferella is E5-16:OAc. This hexadecenyl acetate is a novel moth sex pheromone component.

경북의 자두 주요 생산지인 경산시와 의성군내 8개의 자두과원을 선정하여, 2010년과 2011년에 주요 심식나방류인 복숭아순나방, 복숭아순나방붙이, 복숭아심식나방의 발생상을 성페로몬 트랩을 이용하여 조사하였다. 또한 경산지역에서는 과수원 내부와 외부에서 포획된 개체수 자료를 이용하여 내외부 발생량을 비교하였다. 경산에서 복숭아순나방과 복숭아순나방붙이 발생은 의성보다 약 1주일 정도 빠르게 나타났다. 복숭아순나방은 3월 하순에 발생을 시작하여, 4월 하순 최성기를 이루었으며, 이후 세대는 6월 중순, 7월 중순, 8월 중순에 성충 최성기를 보였다. 반면 복숭아심식나방은 두 지역간의 차이가 없었으며, 6월 상순에 발생을 시작하여 9월 중순까지 2-3회의 발생 최성기를 보였다. 복숭아심식나방의 경우 2010년은 전형적인 2회 최성기를 보였으나 2011년에는 3회 최성기를 보였다. 연간 복숭아순나방, 복숭아순나방붙이, 복숭아심식나방의 발생량비는 경산지역에서 63-47:16-35:20-18%였고, 의성지역은 51-46:18-13:31-36%로 나타났다. 복숭아순나방과 복숭아순나방붙이의 경우 과수원 내부와 외부에서 포획량의 차이는 없었다.

최근 들어 하우스재배 감귤을 중심으로 발생이 증가하고 있는 귤애가루깍지벌레에 대해 효율적인 발생예찰과 함께 방제시기를 결정하기 위하여 합성 성페로몬을 이용하여 시기별 발생특성을 조사하였다. 귤애가루깍지벌레의 수컷 성충은 합성 성페로몬뿐만 아니라 색에도 반응하였다. 성페로몬의 농도는 5.0 mg까지 농도가 증가할수록 유인력이 높았다. 귤애가루깍지벌레 수컷 성충은 연 4회 발생되고 있었으며, 월동 후 수컷 성충이 처음 발생되기 시작하는 시기는 4월 중하순이었다. 수컷 성충발생이 가장 많은 시기는 제 1세대 발생기인 7월 상중순이었다. 성페로몬트 랩에 유인된 귤애가루깍지벌레 수컷 성충은 한 세대 내에서 3회의 발생성기가 나타났으며, 이는 귤애가루깍지벌레가 1~3령 약충으로 월동하기 때문인 것으로 추정되었다. 트랩에 유인된 수컷 성충의 밀도와 나무의 가지당 평균밀도와는 높은 정(+)의 상관을 갖고 있었다. 수컷 성충의 최대 유인시기를 이용하여 1령 약충 발생 시기를 추정할 수 있었으며, 그 적산온도는 산란전 기간과 비슷한 350DD이었다.

This study was conducted to see the initial increasing time of population of citrus leafminer(CLM) in spring and early summer seasons in 2007-2011. CLM adults were monitored using sex pheromone trap which is mainly composed of Z,Z-7, 11-hexadecadienal in the 4, 8 and 5 orchards of the east, south and north part in jeju island, respectively during 2007-2011 seasons. The traps and lures were changed at 10 days and 1 month interval, respectively. Though CLMs were captured on traps from March or April, it was 17, 28 and 28 May in 2007, 16, 26 May and 5 June in 2008, 8, 26 and 26 May in 2009, 27 May, 7 and 7 June in 2010 and 25 May, 2, 2 June in the south, east and north part of Jeju Island, respectively that the CLM population rapidly began to increase toward peaking. The accumulated temperature except less than developmental zero, 12.1 Ujiye T 1990). from January was 248.0, 270.5 and 289.7 in 2007, 193.2, 203.8 and 261.4 in 2008, 202.0, 245.5 and 259.9 in 2009, 261.4, 192.2 and 268.3 in 2010 and 243.4, 246.3 and 252.4 in 2011 at that time. After May overwintered CLM adults are little possible to be counted to the number of captured CLM because adult longevity is about 100 in degree days (Lim 2006). CLM populations in the south part of Jeju Island in first CLM adults increasing periods during 2007-2011 is higher than other parts.

The box tree pyralid, Glyphodes perspectalis, was reared on an artificial diet based on dried powder of box tree leaves, Buxus microphylla var. koreana, and INSECTA F-II (Nihon Nosan Co., Ltd) at a ratio of 3:7 (w:w) and analyzed in their developmental characteristics from 1st to 4th generation. The developmental period of larva reared on artificial diet was longer than that of larva reared on host plant. Significant difference in larval period was also observed between 1st and other generation groups. Pupation rate was the highest in 4th generation followed by 3rd, 2nd, 1st and host plant group. Significant difference in pupal period was noted between 1st and 3rd generation group. Emergence rate was the highest in 4th generation group followed by 3rd, host plant, 1st and 2nd generation group. There is a significant difference in male and female size between 1st and 4th generation group. Our result indicated that stable rearing of box tree pyralid through the year is possible by artificial diet, and the best adaptation to artificial diet was achieved at 4th generation. The pheromone components of this moth have been identified as (Z)-11-hexadecenal (Z11-16:Ald), (E)-11-hexadecenal (E11-16:Ald), and (Z)-11-hexadecenol (Z-11-16:OH). We did several field tests, and found that there was a difference between Korea and Japan population.

The sex pheromone of Synanthedon bicingulata (Staudinger), a major pest of peach trees in many regions of northeast Asia, was identified. Two major components from the pheromone gland extracts of female moths are E3,Z13-18:OAc and Z3,Z13-18:OAc, and the average ratio of these components is about 4:6, respectively. In addition to the major components, four minor components, Z13-18:OAc, E2,Z13-18:OAc, E3,Z13-18:OH, and Z3,Z13-18:OH also were identified from pheromone gland extracts. Field tests showed that E3,Z13-18:OAc and Z3,Z13-18:OAc are essential for attraction of male S. bicingulata moths, and males are optimally attracted to the blend ratio found in pheromone gland extracts of conspecific females. Addition of the minor glandular components (Z13-18:OAc, E2,Z13-18:OAc, E3,Z13-18:OH, and Z3,Z13-18:OH) did not affect captures of males to the primary binary blend. Thus, the blend of E3,Z13-18:OAc and Z3,Z13-18:OAc at the natural ratio can be used for monitoring populations of this species.

In this study, we reared box tree pyralid, Glyphodes perspectalis, with artificial diet and identified sex pheromone. Insect F-II and leaf of box tree were used to make artificial diet. Box tree pyralid, Glyphodes perspectalis, is the most serious pest of box tree in Korea, and was recently introduced into Europe. The pheromone components of this moth have been identified as (Z)-11-hexadecenal (Z11-16:Ald), (E)-11-hexadecenal (E11-16:Ald), and (Z)-11-hexadecenol (Z-11- 16:OH) in Japan. In this study, we identified pheromone components of this species by using GC and GC-MS. Three same components, such as (Z)-11- hexadecenal (Z11-16:Ald), (E)-11-hexdecenal (E11-16:Ald), and (Z)-11-hexadecenol (Z-11-16:OH) were also indentified in Korean population, but there was a little difference in ratios. The ratios of these three compounds of Japanese population and Korean population were 5:1.25:1, and 5:0.96:0.2, respectively. The ratios of Z and E-11-hexadecenal were similar, but the ratio of (Z)-11-hexadecenol was lower compared to Japanese population. In field bioassay, (Z)-11-hexadecenal or (E)-11-hexadecenal alone was not attractive to males, but the mixture of Z11-16:Ald and E11-16:Ald was attractive to males. The most effective ratios of Z11-16:Ald and E11-16:Ald was 5:1. Other ratios such as 1:1 and 1:5 (Z:E) was not attractive to males. The attractiveness of Z:11-16:OH will be tested in near future.

Previous studies indicated that Matsumuraeses phaseoli and M. falcana (Lepidoptera: Tortricidae) are separate species since a few differences were observed in genitalia morphology and female sex pheromone composition. A clear difference was detected in the DNA sequences of cytochrome oxidase I of the two species separately collected in different plants and regions. A hybridization test also showed that a post-zygotic reproductive isolation occurred between the species. In field monitoring, however, both species have been caught simultaneously and together in the separate sex pheromone traps installed for the two species around neighboring soybean and red bean fields. Molecular marker-assisted identification with several adults sampled from the trapped insects showed that only ca. 40% of M. phaseoli adults identified as the species by genitalia morphology was the M. phaseoli, while ca. 97% of M. falcana adults identified as the species was the M. falcana. The result indicated that the observation of genitalia did not make a decisive criterion for classification of the insects. Conclusively, it suggested that the sex pheromones of the two species should be studied more precisely although there is a possibility that the two species are hybridized in fields as in laboratory, and speciation is under process.

Synanthedon haitangvora (Lepidoptera: Sesiidae) is an economically important pest of apple trees in Korea and China. We report here the identification of compounds in pheromone gland extracts of female S. haitangvora, and the male responses to blends of different components and ratios in the field. Two major components from pheromone gland extracts of S. haitangvora females were identified as Z3,Z13-18:OAc and E2,Z13-18:OAc, and the average ratio of these components was about 1:1. Seven minor components, Z9-16:OAc, Z11-16:OAc, Z9-18:OAc, Z13-18:OAc, E3,Z13-18:OAc, Z3,Z13-18:OH, and E2,Z13-18:OH, also were identified from gland extracts. Field tests showed that male S. haitangvora were attracted to Z3,Z13-18:OAc alone, but the maximum number of males was attracted to the binary blend of Z3,Z13-18:OAc and E2,Z13-18:OAc mimicking the blend found in female extracts. The addition of minor components to a 1:1 blend of Z3,Z13-18:OAc and E2,Z13-18:OAc did not increase the numbers of moths captured. The only significant effect of minor components was the strong inhibitory effect of adding Z3,Z13-18:OH to the primary binary blend. Increasing doses of the optimum pheromone blend in the lures from 0.1 to 2.0 mg increased trap catches of male S. haitangvora.